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This chapter describes a teacher leadership professional development program for K-12 science teachers constructed through a partnership between a university and five school districts. The development and implementation of the program drew from the literature on teacher leadership, communities of practice, and distributed leadership. The program supports teachers through a two-year fellowship program where they examine their teaching practices, attend professional development workshops, and undertake an independent teacher leadership project in their own schools. The chapter also describes the research conducted by the university to improve the program and shares findings and future implications of this research.